I think that the one who has changed the most Negatronily is a Phooney because when the Skreeps were first in Vern Hollow Phooney was an Optimist Prime because he was all happy go lucky and didn't take his job seriously and just messed around with Milo. But after he captured the ooman bings and got his arm ripped off by Xafuun he has been all biness like when Grut wanted to give up Phooney went all spider monkey on him and kept kicking him in the booty!

I think that Joe has changed the most because at first in Vern Hollow he was all shady and Daddy Cool. When Jack and Isadora first found him on Arboria they didnt know if the could trust him but now they know they can.